London's Southwark Crown Court was told Mazhar Majeed, 36, was recorded discussing deliberately losing last summer's game with a contact in India.
Pakistan eventually beat England by four wickets at the Oval in London.
Cricketers Salman Butt and Mohammad Asif are accused of fixing parts of a Lord's Test match, which they deny.
Prosecutors allege that Mr Majeed, from Croydon, south London, conspired with Mr Butt, 27, and Mr Asif, 28, to fix parts of the Lord's Test between England and Pakistan last August.
Meanwhile, testimony from earlier in the trial which suggested Australian cricketers were the world's biggest match-fixers has been rejected by Australian cricket's governing body.
London's Southwark Crown Court heard on Monday that Mr Majeed had said Australians were "the biggest" when it came to rigging games.
But Cricket Australia said the claims were "baseless and outlandish". (BBC)